Soils were collected from 15 randomly selected subplots in each plot in early August 2007 and 2009. For bulk density, one soil core (0–10 cm depth, 137 cm3) was collected from the center of each subplot using a bulk density sampler; litter was first removed from the soil surface. For soil chemistry, cores (0–10 cm depth, 35 cm3) were taken with a tube-type sampler (Oakfield Apparatus, Oakfield, Wisconsin, USA). After removing surface litter, cores were extracted from two points adjacent to each of the four vegetation quadrats; the eight subsamples per subplot were composited as they were collected. On the same day, samples were set out to air dry (,258C) for ;72 h.